BioStor
 | Evolutionary relationships, osteology, and zoogeography of leptodactyloid frogs J D Lynch (1971) Miscellaneous Publications of the University of Kansas Museum of Natural History 53: 1-238 |
 | The frog genus Leptodactylus in Ecuador W R Heyer and J A Peters (1971) Proceedings of The Biological Society of Washington 84: 163-170 |