| Frogs of Columbia Doris M Cochran and Coleman J Goin (1970) Bull U S National Museum 288: 1-655 | |
| Leptodactylid frogs of the genus Eleutherodactylus in the Andes of northern Ecuador and adjacent Colombia John D Lynch (1981) University of Kansas Natural History Museum Miscellaneous Publication 72: 1-46 | |
| Frogs of the genus Eleutherodactylus (Leptodactylidae) in western Ecuador: systematics, ecology, and biogeography John D Lynch and William E Duellman (1997) University of Kansas Museum of Natural History Special Publication 23: 1-236 |